PIOCHE SHALE FAUNAL LIST
2003
Compiled by Linda B. McCollum
Geology Department, Eastern Washington University

This faunal list is restricted to the Pioche Shale exposed in Lincoln County, Nevada, and was compiled from both published and unpublished sources.

TRILOBITES:

Grassy Spring Member:
Albertella fritzi Eddy and McCollum, 1998
Albertella highlandensis Eddy and McCollum, 1998
Albertella schenki Resser, 1945
Albertellina aspinosa Palmer, 1979
Albertelloides kitai Eddy and McCollum, 1998
Albertelloides rectimarginatus Palmer, 1979
Albertelloides cf. rectimarginatus Palmer, 1979
Amecephalites sundbergi Eddy and McCollum, 1998
Delamarina brevispinaspis Eddy and McCollum, 1998
Kootenia cf. crassinucha Fritz, 1968
Mexicaspis cf. difuntosensis Lochman, 1948
Mexicella granulata Eddy and McCollum, 1998
Mexicella mexicana Lochman, 1948
Mexicella stator (Walcott, 1916)
Nyella granosa (Resser, 1939)
Nyella immoderata Palmer, 1979
Olenoides maladensis Resser, 1939
Onchocephalites? sp.
Pachyaspis sp. undet.
Pagetia resseri Kobayashi, 1943
Panacus palmeri Eddy and McCollum, 1998
Paralbertella sp. undet.
Plagiura minor Palmer, 1979
Poliella denticulata Rasetti, 1951
Poliella germana (Resser, 1939)
Poliella cf. germana (Resser, 1939)
Ptarmiganoides hexacantha Palmer, 1979
Ptarmiganoides propinqua (Resser, 1939)
Ptarmiganoides? sp.
Ursinella major Eddy and McCollum, 1998
Volocephalina connexa Palmer, 1979
Volocephalina variosa Eddy and McCollum, 1998
Zacanthoides? sp.
Corynexochid genus and species unidentified
 

Log Cabin Member
Amecephalus sp. unident.
Eokochaspis? cabinensis Sundberg and McCollum, 2003
Fieldaspis celer (Walcott, 1917)
Fieldaspis superba Rasetti, 1951
Kochaspis liliana (Walcott, 1886)
Kochiella augusta (Walcott, 1886)
Kochiella brevaspis Sundberg and McCollum, 2002
Kochiella maxeyi Rasetti, 1951
Kochiellina janglensis Palmer, 1979
Hadrocephalites lyndonensis Sundberg and McCollum, 2002
Hadrocephalites rhytidodes Sundberg and McCollum, 2002
Hadrocephalites sp. A Sundberg and McCollum, 2002
Hadrocephalites sp. B Sundberg and McCollum, 2002
Mexicella mexicana Lochman, 1948
Nyella? plana Sundberg and McCollum, 2003
Oryctocephalus primus Walcott, 1886
Oryctocephalus sp. unident.
Parapoulsenia cf. lata Rasetti, 1957
Piochella peasleensis Sundberg and McCollum, 2003
Plagiura extensa Palmer, 1979
Poliella ovala Sundberg and McCollum, 2003
Poliella cf. prima (Walcott, 1908)
Poliellaites gloriosa Sundberg and McCollum, 2003
Stephenaspis highlandensis Sundberg and McCollum, 2003
Syspacephalus sp. unident.
 

Susan Duster Limestone Member
Amecephalus arrojosensis (Lochman, 1952)
Caborcella reducta Palmer, 1979
Fieldaspis bilobata Rasetti, 1951
Hadrocephalites rhytidodes Sundberg and McCollum, 2002
Kochiella brevaspis Sundberg and McCollum, 2002
Kochiella rasettii Sundberg and McCollum, 2002
Mexicella robusta Sundberg and McCollum, 2000
Nyella? plana Sundberg and McCollum, 2003
Nyella rara (Rasetti, 1951)
Onchocephalites sp. unident.
Oryctocephalus n. sp.
Poliella denticulata Rasetti, 1951
Poliella lomataspis Palmer, 1979
Syspacephalus crassus Rasetti, 1951
 

Comet Shale Member
Amecephalus arrojosensis (Lochman, 1952)
Amecephalus sp. A Sundberg and McCollum, 2000
Eokochaspis delamarensis Sundberg and McCollum, 2000
Eokochaspis longspina Sundberg and McCollum, 2000
Eokochaspis nodosa Sundberg and McCollum, 2000
Eokochaspis piochensis (Palmer, 1979)
Kochina? walcotti Sundberg and McCollum, 2000
Mexicella antelopea Sundberg and McCollum, 2000
Mexicella robusta Sundberg and McCollum, 2000
Oryctocephalites rasettii Sundberg and McCollum, 1997

Combined Metals Member
Bathynotus holopygus (Hall, 1859)
Bolbolenellus brevispinus Palmer, 1998
Crassifimbra walcotti (Resser, 1937)
Eokochapis metalaspis Sundberg and McCollum, 2000
Nephrolenellus geniculatus Palmer, 1998
Nephrolenellus multinodus Palmer, 1979
Olenellus chiefensis Palmer, 1998
Olenellus fowleri Palmer, 1998
Olenellus gilberti (Meek, 1874)
Olenellus howelli (Meek, 1874)
Olenellus terminatus Palmer, 1998
Oryctocephalites palmeri Sundberg and McCollum, 1997
Zacanthopsis levis (Walcott, 1886)
Zacanthopsis n. sp.

Delamar Member
Bristolia bristolensis (Resser, 1928)
Bristolia harringtoni Lieberman, 1999
Bristolia insolens (Resser, 1928)
Bristolia mohavensis (Resser, 1928)
Mesonacis n. sp.
?Nephrolenellus n. sp.
Olenellus clarki (Resser, 1928)
Olenellus fremonti Walcott, 1910
Olenellus nevadensis (Walcott, 1910)
Peachella iddingsi (Walcott, 1884)
Peachella n. sp.

Lagerstatten:
Anomalocaris pennsylvanica
Anomalocaris cf. saron
Canadaspis perfecta
Ottoia sp. unident.
?Perspicaris dilatus
Tuzoia guntheri
Tuzoia nitida
Tuzoia polleni
 

INARTICULATE BRACHIOPODS
Acrothele spp.
Acrothyra minor Walcott, 1905
Aphelotreta minuta Rowell, 1980
Dictyonina pannula (White, 1874)
Eothele spurri (Walcott, 1908)
Hadrotreta primaaea (Walcott, 1902)
Micromitra sp.

ARTICULATE BRACHIOPODS
Wymenella highlandensis (Walcott, 1886)

HELCIONELLOIDA
Igorella insulcata (Rasetti, 1957)
Helcionella? carinata Rasetti, 1957
Latouchella arguta (Resser, 1939)
 

STENOTHECOIDA
Stenothecoides elongata (Walcott, 1884)
 
 

ICHNOFAUNA

CHISHOLM SHALE
Monocraterion
Paleoscolecida
Planolites
Psammichnites "sardensis"
Sericichnus mureroensis
Skolithos

PIOCHE SHALE (26g., 29 sp.)

Grassy Spring Member (21 g., 24 sp.)
Aulichnites
Bergaueria Prantl, 1946
Bifungites Desio, 1940
Cochlichnus Hitchcock, 1858
Cruziana D'Orbigny, 1842
Diplichnites Dawson, 1873
Diplocraterion parallelum Torell, 1870
Margaritichnus Bandel, 1973
Monocraterion Torell, 1870
Monomorphichnus lineatus Crimes, 1970
Neonereites biseriales Seilacher, 1960
N. uniseriales
Phycodes palmatum Richter, 1850
Planolites beverleyensis Nicholson, 1873
P. montanus
P. terraenovae
Rhizocorallium Zenker, 1836
Rusophycus Hall, 1852
Scolicia De Quatrefages, 1849
Skolithos linearis Haldemann, 1840
Taenidium serpentinum Heer, 1877
?Taphrhelminthopsis Sacco, 1888
Teichichnus rectus Seilacher, 1955
Treptichnus bifurcus Miller, 1889

Log Cabin Member (18 g., 19 sp.)
Arenicolites Salter, 1857
Bergaueria
Bifungites
Cruziana
Cylindrichnus concentricus Toots, in Howard, 1966
Dimorphichnus obliquus Seilacher, 1955
Diplichnites
Diplocraterion
Monocraterion
Monomorphichnus lineatus Crimes, 1970
Monomorphichnus multilineatus
Phycodes
Planolites terraenovae
Psammichnites "sardensis" Torell, 1870
Rusophycus
Scolicia prisca
Sericichnus
Skolithos
Teichichnus rectus Seilacher, 1955

Comet Shale Member (7 g. and sp.)
Bifungites
Cylindrichnus concentricus
Diplichnites
Monomorphichnus
Planolites montanus
Rusophycus
Sericichnus

Combined Metals Member (8 g. and sp.)
Arenicolites
Cochlichnus
Cruziana
Monocraterion
Planolites montanus
Sericichnus
Skolithos
Taphrhelminithopsis

Delamar Member (6 g. and sp.)
Monomorphichnus lineatus Crimes, 1970
Planolites beverleyensis Nicholson, 1873
Phycodes
Sericichnus
Skolithos
Teichichnus
 
 

ZABRISKIE QUARTZITE
Diplocraterion
Monocraterion
Planolites
Rusophycus cantabricus (bonnerensis)
Skolithos
 
 

References

Eddy, J.D., and McCollum, L.B., 1998, Early Middle Cambrian Albertella Biozone trilobites of the Pioche Shale, southeastern Nevada. Journal of Paleontology, v. 72, p. 864-887.

Merriam, C.W., 1964, Cambrian rocks of the Pioche mining district, Nevada. U.S. Geological Survey Professional Paper 469, 59 p.

Palmer, A.R., 1998, Terminal Early Cambrian extinction of the Olenellina: Documentation from the Pioche Formation, Nevada. Journal of Paleontology, v. 72, p. 650-672.

Rowell, A.J., 1980, Inarticulate brachiopods of the Lower and Middle Cambrian Pioche Shale of the Pioche District, Nevada. University of Kansas Paleontological Contribution, Paper 98, 26 p.

Sundberg, F.A., and McCollum, L.B., 1997, Oryctocephalids (Corynexochida: Trilobita) of the Lower-Middle Cambrian boundary interval from California and Nevada. Journal of Paleontology, v. 71, p. 1065-1090.

Sundberg, F.A., and McCollum, L.B., 2000, Ptychopariid trilobites from the Lower-Middle Cambrian boundary interval, Pioche Shale, southeastern Nevada. Journal of Paleontology, v. 74, p. 604-630.

Sundberg, F.A., and McCollum, L.B., 2002, Kochiella Poulsen, 1927, and Hadrocephalites new (Ptychopariida, Trilobita) from the early Middle Cambrian of western Laurentia. Journal of Paleontology, v. 76, p. 76-94.

Sundberg, F.A., and McCollum, L.B., 2003, Trilobites of the lower Middle Cambrian Poliella denticulata Biozone (new) of southeastern Nevada.  Journal of Paleontology, v. 77, p. 333-361.

Home